Bacterial and archaeal symbioses with protists
https://www.cell.com/current-biology/fulltext/S0960-9822(21)00747-8
all of the data Garth has about extremophiles being used to enhance or weaponize microbes, the original gain of function, also points to Protist Endosymbionts, not so much Archaea Carboxysome, like i previous thought, but its difficult to find confirmation.
but as you can see here, we still have a long road to understanding the kind of chimera possibilities.
RE: Protonation Cation/Anion